# Cooperation
> **Machine-Evolution Nexus:** [[articles/Digital Darwinism and the Invisible World of Machine Evolution|Digital Darwinism and the Invisible World of Machine Evolution]] places this node within the history and governance of substrate-independent evolution.
**Domain:** Evolution / Social Systems
**Doc Type:** Canonical Wiki Node
**Maturity:** Developed
## Definition
Cooperation occurs when interaction produces benefits for others or a collective, often at a cost or risk to the contributor.
## Nexus Context
Digital evolution allows cooperation to arise and be tested under known relatedness, spatial and resource conditions. It is one possible response to selection, not evolution’s default moral direction.
## Evidence Boundary
Functional cooperation does not establish altruistic intention or subjective solidarity. Governance must distinguish behavioral outcome from inner motive.
## Relationships
[[wiki/Division of Labor|Division of Labor]], [[wiki/Symbiosis|Symbiosis]], [[wiki/Major Evolutionary Transitions|Major Evolutionary Transitions]], [[wiki/Avida|Avida]]
## Sources / Provenance
- Evolution of cooperation literature; Avida experiments.